Checklist of herpetofauna
listed in the CITES appendices
and in EC Regulation No. 338/97
10th Edition
2005
compiled by
UNEP-WCMC
© JNCC 2005
The JNCC is the forum through which the three country conservation agencies - the
Countryside Council for Wales, English Nature and Scottish Natural Heritage - deliver
their statutory responsibilities for Great Britain as a whole, and internationally. These
responsibilities contribute to sustaining and enriching biological diversity, enhancing
geological features and sustaining natural systems. As well as a source of advice and
knowledge for the public, JNCC is the Government's wildlife adviser, providing guidance
on the development of policies for, or affecting, nature conservation in Great Britain or
internationally.
Published by: Joint Nature Conservation Committee
Copyright: 2005 Joint Nature Conservation Committee
ISBN: 1st edition published 1979 ISBN 0-86139-075-X
2nd edition published 1981 ISBN 0-86139-095-4
3rd edition published 1983 ISBN 0-86139-224-8
4th edition published 1988 ISBN 0-86139-465-8
5th edition published 1993 ISBN 1-873701-46-2
6th edition published 1995 ISSN 0963-8091
7th edition published 1999 ISSN 0963-8091
8th edition published 2001 ISSN 0963-8091
9th edition published 2003 ISSN 0963-8091
10th edition published 2005 ISSN 0963-8091
Citation: UNEP-WCMC (2005). Checklist of herpetofauna listed in the CITES
appendices and in EC Regulation 338/97. 10th edition. JNCC Report No.
378.
Further copies of this report are available from:
CITES Unit
Joint Nature Conservation Committee
Monkstone House
City Road
Peterborough PE1 1JY
United Kingdom
Tel: +44 1733 562626
Fax: +44 1733 555948
This document can also be downloaded from:
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Prepared under contract from the Joint Nature Conservation Committee by UNEPWCMC.
The UNEP World Conservation Monitoring Centre was established in 2000 as the
world biodiversity information and assessment centre of the United Nations Environment
Programme. The roots of the organization go back to 1979, when it was founded as the
IUCN Conservation Monitoring Centre. In 1988 the World Conservation Monitoring Centre
was created jointly by IUCN, WWF International and UNEP. The financial support and
guidance of these organizations in the Centre’s formative years is gratefully
acknowledged.
The designations of geographical entities in this report, and the presentation of the
material, do not imply the expression of any opinion whatsoever on the part of JNCC or
WCMC concerning the legal status of any country, territory, or area, or of its authorities,
or concerning the delimitation of its frontiers or boundaries.
Contents
Acknowledgements.................... i
Introduction .............................. i
Explanatory notes..................... ii
Introductory References............v
CHORDATA...................................... 1
REPTILIA ..................................... 1
TESTUDINES .............................. 1
DERMATEMYDIDAE........................ 1
PLATYSTERNIDAE ......................... 1
EMYDIDAE ................................... 1
TESTUDINIDAE............................. 7
CHELONIIDAE............................. 12
DERMOCHELYIDAE...................... 15
CARETTOCHELYIDAE ................... 15
TRIONYCHIDAE........................... 15
PELOMEDUSIDAE ........................ 17
CHELIDAE .................................. 19
CROCODYLIA............................ 19
ALLIGATORIDAE ......................... 19
CROCODYLIDAE.......................... 21
GAVIALIDAE............................... 23
RHYNCHOCEPHALIA................. 23
SPHENODONTIDAE...................... 23
SAURIA .................................... 23
GEKKONIDAE ............................. 23
AGAMIDAE ................................. 30
CHAMAELEONIDAE...................... 31
IGUANIDAE................................ 44
LACERTIDAE............................... 46
CORDYLIDAE.............................. 46
GERRHOSAURIDAE...................... 50
TEIIDAE..................................... 50
SCINCIDAE ................................ 51
XENOSAURIDAE.......................... 51
HELODERMATIDAE ...................... 51
VARANIDAE................................ 52
SERPENTES .............................. 58
LOXOCEMIDAE ........................... 58
PYTHONIDAE.............................. 58
BOIDAE ..................................... 62
BOLYERIIDAE ............................. 67
TROPIDOPHIIDAE ....................... 67
COLUBRIDAE.............................. 69
ELAPIDAE .................................. 71
HYDROPHIIDAE .......................... 73
VIPERIDAE................................. 73
AMPHIBIA.................................. 74
ANURA ..................................... 74
BUFONIDAE................................ 74
DENDROBATIDAE........................ 75
MANTELLIDAE ............................ 81
MICROHYLIDAE .......................... 82
MYOBATRACHIDAE ...................... 83
RANIDAE.................................... 83
CAUDATA.................................. 83
AMBYSTOMIDAE.......................... 83
CRYPTOBRANCHIDAE................... 84
References.............................. 85
Index .................................... 122
i
Acknowledgements
This checklist was compiled by UNEP-WCMC under contract with the Joint Nature
Conservation Committee.
The volume builds on earlier editions, and the numerous contributors to those editions
are acknowledged. UNEP-WCMC staff involved in the production of this volume include
Tim Inskipp, Sarah Ferriss, James O’Carroll and Tobias Garstecki. The project was
supervised by Gerardo Fragoso.
Vin Fleming, Alison Littlewood and Nichola Burnett of the JNCC CITES Unit are thanked
for providing advice and guidance throughout.
Introduction
In April 1991, the Nature Conservancy
Council for England (English Nature),
Countryside Council for Wales and
Scottish Natural Heritage acting together
through the Joint Nature Conservation
Committee were appointed by the
Secretary of State for the Environment
as the United Kingdom's Scientific
Authority for Animals under the
Convention on International Trade in
Endangered Species of Wild Fauna and
Flora (CITES). CITES regulates
international trade in wild animals and
plants and in products derived from
them, to help to ensure their
conservation on a worldwide scale.
The intention of this publication is to
provide a list of the species and
subspecies of mammal included in the
CITES Appendices, together with their
conservation status category in the 2004
IUCN Red List of threatened species
(IUCN, 2004) and the appropriate Annex
in EC Regulation 338/97 (and
subsequent updates).
This revised edition incorporates
additions and amendments to the CITES
appendices up to and including those
made at the 13th Conference of the
Parties in Thailand in October 2004
(effective from 12 January 2005) and
those outlined in CITES Notification
2004/074 and Notification 2005/029.
The most recent EC Regulation to amend
the Annexes is 1332/2005, effective
from 12 August 2005.
For background material on the rationale
of the IUCN threat categories, readers
are referred to IUCN (2004); for
background on the workings of CITES,
recommended works are Favre (1989)
and Wijnstekers (2003). For information
on the implementation of CITES in the
EU, see European Commission (2005).
For information by country on the
diversity and status of herpetofauna and
other taxonomic groups, and for a
general review of biodiversity, readers
are referred to Groombridge and Jenkins
(2002).
In a publication of this nature, it is
inevitable that users will discover entries
that need correcting or updating. The
publishers and UNEP-WCMC would be
grateful to receive details of those
entries so that the necessary changes
can be made in the database from which
this document is produced.
The database is continually updated and
the downloaded version of this document
may contain more recent information
than the original printed publication.
ii
Explanatory notes
Each species is typically represented by
a block of text including scientific name
(with alternatives), common names in
the three official CITES languages
(where available), geographic range,
CITES Appendix, EC Regulation 338/97
Annex, and Red List category. Where a
species has component populations or
formal subspecies which are treated
differently by CITES, the species entry is
subdivided appropriately.
Scientific names
For each taxon, the scientific name is
given first (as listed in the CITES
appendices), with the most frequently
used synonyms listed beneath the
scientific name.
The sequence of families within orders
conforms with the CITES Appendices
(thus the amphibian families are in
alphabetical order while the reptile
families follow a conventional systematic
order). The genera and species are in
alphabetical order within each family.
The taxonomic sources adopted by
CITES are given below.
Reptiles:
Testudines (Wermuth and Mertens,
1996; Karl & Tichy, 1999; McCord et
al., 2000; McCord & Pritchard, 2002;
Perälä, 2001; Webb, 2002).
Crocodylia (Wermuth and Mertens,
1996).
Rhynchocephalia (Wermuth and Mertens,
1996).
Sauria (Pough et al, 1998 for
delimitation of families within the
Sauria)
Phelsuma (Hallmann et al., 1997;
Nussbaum et al., 2000; Rösler et al.,
2001)
Uromastyx (Wilms, 2001)
Chamaeleonidae (Klaver and Böhme,
1997, except for the recognition of
Calumma andringitaensis, C.
guillaumeti, C. hilleniusi &
C. marojezensis as valid species; also
Andreone et al., 2001; Böhme, 1997;
Jesu et al., 1999; Tilbury, 1998).
Cordylidae (Broadley, 2002).
Crocodilurus (Massary & Hoogmoed,
2001).
Tupinambis (Avila Pires, 1995; Cei,
1993; Manzani & Abe, 1997; Manzani
& Abe, 2002; Colli et al. 1998).
Varanidae (Böhme, 2003; Jacobs, 2003)
Boidae (McDiarmid et al., 1999 except
for the retention of the genera
Acrantophis, Sanzinia, Calabaria &
Lichanura and the recognition of
Epicrates maurus as a valid species;
also Dirksen, 2002).
Bolyeriidae (McDiarmid et al., 1999).
Loxocemidae (McDiarmid et al., 1999).
Pythonidae (McDiarmid et al., 1999;
Harvey et al., 2000; Keogh et al.,
2001).
Tropidophiidae (McDiarmid et al., 1999;
Hedges et al. 1999; Hedges &
Garrido, 1999; Hedges & Garrido,
2002; Hedges et al., 2001).
Naja (Wüster, 1996; Slowinski & Wüster,
2000).
Viperidae (McDiarmid et al., 1999).
No standard references have been
adopted by CITES for
Hoplodactylus spp., Naultinus spp.,
Uroplatus spp., Brachylophus spp.,
Conolophus spp., Cyclura spp., Iguana
spp., Dracaena spp. & Heloderma spp.
Amphibians: Frost (2002)
Common names
Common names (in English, French and
Spanish where available) appear on the
line immediately following the scientific
name and synonyms. However, few of
the listed taxa have regularly or
universally used common names. In
some cases, identical trade names
appear to be used for different taxa.
Secondary English common names have
been included wherever this was
considered useful, including non-English
names commonly used by English
speakers.
Spanish and French names have been
taken from the CITES Identification
Manual Series and other sources.
Other information
The three columns headed CITES, EC
Reg. and RL list the following information
for each taxon.
iii
CITES
I, II or III in this column refers to the
appendix in which the taxon is listed
under the Convention on International
Trade in Endangered Species of Wild
Fauna and Flora. NC = non-CITES
The two-letter abbreviations following
Appendix III entries denote the countries
whose governments have placed the
taxon in this appendix, as follows:
CN China
GH Ghana
HN Honduras
IN India
NZ New Zealand
EC Reg.
The letters in this column refer to the
Annex of EC Regulation 338/97 in which
the taxon is listed. NR= non Regulation.
RL
Threat categories follow those given in
the 2004 IUCN Red List (IUCN, 2004)
and are as follows:
EX Extinct
EW Extinct in the Wild
CR Critically Endangered
EN Endangered
VU Vulnerable
LR Lower Risk
LR/cd Lower Risk/Conservation
dependent
LR/nt Lower Risk/Near threatened
LR/lc Lower Risk/Least concern
DD Data Deficient
NE Not Evaluated
Note that the current Red List includes
assessments using both the 1994 and
the 2001 Categories and Criteria.
Geographical range
The geographical range of each taxon is
generally given in terms of political units
arranged alphabetically. Small island
dependencies are also listed
alphabetically.
Place-names and names of countries
follow the Times Atlas (2003) and United
Nations Cartographic Section (2004).
It should be noted that, when a species
is listed as occurring within a given area
or range state, it may not occur
throughout that country or area, and
may be confined to only one or a few
localities. This is particularly true for
migratory species.
Without surveying the range of each
taxon in the field, it is necessary to rely
on published records. Many taxonomic
works give the range of a taxon in terms
of broad geographical areas, rather than
of political units. While some countries
have a relatively well-known and
well-reported herpetofauna, some others
do not do so. These factors may
occasionally have resulted in some of the
geographical ranges given here being
incomplete or inaccurate, although every
effort has been made to prevent this.
Distribution notes
A question mark ‘?’ preceding a
geographical range has been used to
indicate that there is some uncertainty in
the occurrence of the species in that
range.
A country where the species is known to
be extinct is denoted by ‘(ex)’, or by
‘(ex?)’ where there is still a small
possibility that the species survives, or
where recent searches have been
unsuccessful.
Range states where the species has been
introduced are denoted by ‘[int]’. Range
states where the species has been reintroduced
are denoted by '[re-int]'.
References
The numbers in the right-hand column
refer to entries in the reference list at
the end of this document. Many of these
are general works relevant to more than
one geographical range or concerned
with particular groups of species. Single
country faunas and more specific
references have generally been inserted
in brackets after the appropriate country
in the listings for geographical range.
iv
Dependent territories
In the text, parent countries of island groups or dependent territories are
omitted. These are listed below:
American Samoa, USA
Anguilla, United Kingdom
Aruba, Netherlands
Azores, Portugal
Bermuda, United Kingdom
Bouvet Island, Norway
British Indian Ocean Territory, United
Kingdom
British Virgin Islands, United
Kingdom
Cayman Islands, United Kingdom
Christmas Island, Australia
Cocos (Keeling) Islands, Australia
Cook Islands, New Zealand
Falkland Islands, United Kingdom
Faeroe Islands, Denmark
Federated States of Micronesia, USA
French Guiana, France
French Polynesia, France
French Southern and Antarctic
Territories, France
Gibraltar, United Kingdom
Greenland, Denmark
Guadeloupe, France
Guam, USA
Heard and Macdonald Islands,
Australia
Marshall Islands, USA
Martinique, France
Mayotte, France
Montserrat, United Kingdom
Netherlands Antilles, Netherlands
New Caledonia, France
Niue, New Zealand
Norfolk Island, Australia
Northern Marianas, USA
Pitcairn Islands, United Kingdom
Puerto Rico, USA
Réunion, France
Saint Helena, United Kingdom
(including its dependencies of
Ascension and Tristan da Cunha)
Saint Pierre and Miquelon, France
Svalbard and Jan Mayen Islands,
Norway
Tokelau, New Zealand
Turks and Caicos Islands, United
Kingdom
Virgin Islands of the United States,
USA
Wallis and Futuna, France
v
Introductory references
Andreone, F., Mattioli, F., Jesu, R. and
Randrianirina, J. E. (2001) Two new
chameleons of the genus Calumma from
north-east Madagascar, with observations on
hemipenial morphology in the Calumma
furcifer group (Reptilia, Squamata,
Chamaeleonidae). Herpetological Journal 11:
53-68.
Avila Pires, T. C. S. 1995. Lizards of
Brazilian Amazonia. Zool. Verh. 299: 706
pp.
Böhme, W. 1997. Eine neue Chamäleon art
aus der Calumma gastrotaenia –
Verwandtschaft Ost-Madagaskars.
Herpetofauna (Weinstadt) 19 (107): 5-
10.
Böhme, W. 2003. Checklist of the living
monitor lizards of the world (family
Varanidae). Zool. Verh. Leiden 341: 4-
43.
Broadley, D. G. 2002. CITES Standard
reference for the species of Cordylus
(Cordylidae, Reptilia) prepared at the
request of the CITES Nomenclature
Committee. CoP12 Inf. 14.
Cei, J. M. 1993. Reptiles del noroeste,
nordeste y este de la Argentina –
herpetofauna de las selvas subtropicales,
puna y pampa. Monografie XIV, Museo
Regionale di Scienze Naturali.
Colli, G. R., Péres, A. K. and da Cunha, H. J.
1998. A new species of Tupinambis
(Squamata: Teiidae) from central Brazil,
with an analysis of morphological and
genetic variation in the genus.
Herpetologica 54: 477-492.
Dirksen, L. 2002. Anakondas. NTV
Wissenschaft.
European Commission 2005. The European
Community and trade in wild fauna and
flora.
http://europa.eu.int/comm/environment/
cites/home_en.htm
Favre, D. S. 1989. International trade in
endangered species: a guide to CITES.
Martinus Nijhoff, Dordrecht.
Frost, D. R., ed. 2002. Amphibian Species of
the World: a taxonomic and geographic
reference.
http://research.amnh.org/herpetology/a
mphibia/index.html as of 23 August 2002
Groombridge, B. and Jenkins, M.D. 2002.
World atlas of biodiversity. Prepared by
the UNEP World Conservation Monitoring
Centre. University of California Press,
Berkeley, USA.
Hallmann, G., Krüger, J. and Trautmann, G.
1997. Faszinierende Taggeckos – Die
Gattung Phelsuma. Natur & Tier-Verlag.
Harvey, M. B., Barker, D. B., Ammerman, L.
K. and Chippindale, P. T. 2000.
Systematics of pythons of the Morelia
amethistina complex (Serpentes: Boidae)
with the description of three new species.
Herpetological Monographs 14: 139-185.
Hedges, B. S., Estrada, A. R. and Diaz, L. M.
1999. New snake (Tropidophis) from
western Cuba. Copeia 1999(2): 376-381.
Hedges, S. B. and Garrido, O. 1999. A new
snake of the genus Tropidophis
(Tropidophiidae) from central Cuba.
Journal of Herpetology 33: 436-441.
Hedges, B. S. and Garrido, O. 2002. A new
snake of the genus Tropidophis
(Tropidophiidae) from eastern Cuba.
Journal of Herpetology 36:157-161.
Hedges, B. S., Garrido, O. and Diaz, L. M.
2001. A new banded snake of the genus
Tropidophis (Tropidophiidae) from northcentral
Cuba. Journal of Herpetology 35:
615-617.
IUCN. 2004. 2004 IUCN Red List of
threatened species. IUCN, Gland,
Switzerland. www.redlist.org
Jacobs, H. J. 2003. A further new emerald
tree monitor lizard of the Varanus
prasinus species group from Waigeo,
West Irian (Squamata: Sauria:
Varanidae). Salamandra 39(2): 65-74.
Jesu, R., Mattioli, F. and Schimenti, G. 1999.
On the discovery of a new large
chameleon inhabiting the limestone
outcrops of western Madagascar: Furcifer
nicosiai sp. nov. (Reptilia,
Chamaeleonidae). Doriana 7(311): 1-14.
Karl, H.-V. and Tichy, G. 1999. Die
Taxonomie von Homopus bergeri
v i
Lindholm 1906 (Testudines:
Testudinoidea). Mauritiana 17: 277-284.
Keogh, J. S., Barker, D. G. and Shine, R.
2001. Heavily exploited but poorly
known: systematics and biogeography of
commercially harvested pythons (Python
curtus group) in Southeast Asia.
Biological Journal of the Linnean Society
73: 113-129.
Klaver, C. J. J. and Böhme, W. 1997.
Chamaeleonidae. Das Tierreich 112: 85
pp.
Manzani, P. R. and Abe, A. S. 1997. A new
species of Tupinambis Daudin, 1802
(Squamata, Teiidae) from central Brazil.
Boletim do Museu Nacional Nov. Ser.
Zool. 382: 1-10.
Manzani, P. R. and Abe, A. S. 2002. A new
species of Tupinambis Daudin, 1803 from
southeastern Brazil (Squamata, Teiidae).
Arquivos do Museu Nacional, Rio de
Janeiro 60(4): 295-302.
Massary, J.-C. de and Hoogmoed, M. 2001.
The valid name for Crocodilurus
lacertinus auctorum (nec Daudin, 1802)
(Squamata: Teiidae). Journal of
Herpetology 35: 353-357.
McCord, W. P., Iverson, J. B., Spinks, P. Q.
and Shaffer, H. B. 2000. A new genus of
Geoemydid turtle from Asia. Hamadryad
25: 86-90.
McCord, W. P. and Pritchard, P. C. H. 2002.
A review of the softshell turtles of the
genus Chitra, with the description of new
taxa from Myanmar and Indonesia (Java).
Hamadryad 27 (1): 11-56.
McDiarmid, R. W., Campbell, J. A. and
Touré, T. A. 1999. Snake Species of the
World. A Taxonomic and Geographic
Reference. Volume 1. The Herpetologists’
League, Washington, DC.
Nussbaum, R. A., Raxworthy, C. J.,
Raselimanana, A. P. and Ramanamanjato,
J. B. 2000. New species of day gecko,
Phelsuma Gray (Reptilia: Squamata:
Gekkonidae), from the Reserve Naturelle
Integrale d'Andohahela, south
Madagascar. Copeia 2000: 763-770.
Perälä, J. 2001. A new species of Testudo
(Testudines: Testudinidae) from the
Middle East, with implications for
conservation. Journal of Herpetology 35:
567-582.
Pough, F. H., Andrews, R. M., Cadle, J. E.,
Crump, M. L., Savitzky, A. H. and Wells,
K. D. 1998. Herpetology.
Rösler , H., Obst, F. J. and Seipp, R. 2001.
Eine neue Taggecko-Art von
Westmadagaskar: Phelsuma hielscheri sp.
n. (Reptilia: Sauria: Gekkonidae).
Zoologische Abhandlungen, Staatliches
Museum für Tierkunde, Dresden 51: 51-
60.
Slowinski, J. B. and Wüster, W. 2000. A new
cobra (Elapidae: Naja) from Myanmar
(Burma). Herpetologica 56: 257-270.
The Times Atlas of the World 2003.
Comprehensive (eleventh) edition. Times
Books, London UK.
Tilbury, C. 1998. Two new chameleons
(Sauria: Chamaeleonidae) from isolated
Afromontane forests in Sudan and
Ethiopia. Bonner Zoologische Beiträge
47: 293-299.
United Nations Cartographic Section, 2004.
List of Territories. Technical Paper 2004.
http://www.un.org/Depts/Cartographic/e
nglish/geoname.pdf
Webb, R. G. 2002. Observations on the
Giant Softshell Turtle, Pelochelys cantorii,
with description of a new species.
Hamadryad 27 (1): 99-107.
Wermuth, H. and Mertens, R. 1996 (reprint).
Schildkröte, Krokodile, Brückenechsen.
Gustav Fischer Verlag, Jena.
Wijnstekers, W. 2003. The evolution of
CITES. Seventh edition. CITES
Secretariat, Lausanne, Switzerland.
Wilms, T. 2001. Dornschwanzagamen:
Lebensweise, Pflege, Zucht. Herpeton
Verlag.
Wüster, W. 1996. Taxonomic change and
toxinology: systematic revisions of the
Asiatic cobras Naja naja species complex.
Toxicon 34: 339-406.
Phylum: CHORDATA
Class: REPTILIA
Order: TESTUDINES
Family: PLATYSTERNIDAE
Platysternon megacephalum Gray, 1831 II B EN
E: Big-headed Turtle
China [3396, 5050]; Hong Kong, China [5227]; Lao People's Democratic Republic [5037]; Myanmar [3396, 10420];
Thailand [3396, 5053, 6262]; Viet Nam [3396]
Family: DERMATEMYDIDAE
Dermatemys mawii Gray, 1847 II B EN 3396
Synonyms: Limnochelone micrura Werner, 1901, Emys berardii Duméril, Bibron & Duméril, 1851, Emys
dermatemys Duméril & Bocourt, 1888, Dermatemys abnormis Cope, 1868, Dermatemys salvinii Gray, 1870
E: Central American River Turtle, F: Dermatemyde de Mawe, Tortue de Tabasco, S: Plana, Tortuga blanca
Belize [3525]; Guatemala; Honduras; Mexico
CITES Identification Manual Reference: A-301.005.001.001
Family: EMYDIDAE
Annamemys annamensis (Siebenrock, 1903) II B
Synonyms: Annamemys groschovskiae Tien, 1957, Annamemys merkleni Bourret, 1939, Mauremys
annamensis (Siebenrock, 1903), Cyclemys annamensis Siebenrock, 1903
E: Annam Leaf Turtle, Annam Pond Turtle, F: Emyde d'Annam
Viet Nam [3396, 5173]
Batagur baska (Gray, 1831) I A CR 3396, 5226
Synonyms: Tetronyx longicollis Lesson, 1832, Tetraonyx lessonii Duméril & Bibron, 1835, Emys baska Gray,
1831, Emys batagur Gray, 1831, Emys tetraonyx Temminck & Schlegel, 1833
E: Batagur, Common Batagur, Four-toed Terrapin, River Terrapin, F: Batagur malais, Émyde fluviale
indienne, S: Galápago Batagur, Galápago indio
Bangladesh [3396, 5228, 5236]; Cambodia [3396, 5143, 25563]; India [3263, 3264, 3396, 5175]; Indonesia [3396, 5094]:
Sumatera [5085, 5094]; Malaysia [3396, 5095]: Peninsular Malaysia; Myanmar [3396, 5084, 10420]; ?Singapore;
Thailand (ex) [3396, 5053, 5081, 6262]; Viet Nam (ex)
CITES Identification Manual Reference: A-301.007.002.001
Callagur borneoensis
(Schlegel & S. Müller, 1844) II B CR 4988, 5226
Synonyms: Callagur picta (Gray, 1862), Batagur picta Gray, 1862, Emys borneoensis Schlegel & S. Müller,
1844, Kachuga major Gray, 1873, Kachuga brookei Bartlett, 1895
E: Painted Batagur, Painted Terrapin, Saw-jawed Turtle, Three-striped Batagur, F: Émyde peinte de Bornéo,
S: Galápago pintado
?Brunei Darussalam; Indonesia [5094]: Kalimantan [3396, 5094], Sumatera [3396, 5085, 5094]; Malaysia [3396,
5095] : Peninsular Malaysia [3396], Sarawak [3396]; Thailand [3396, 5053, 6262]
Chinemys megalocephala Fang, 1934 III CN C EN 10455
E: Big-headed Pond Turtle
China [5050]
Chinemys nigricans (Gray, 1834) III CN C EN
Synonyms: Chinemys kwangtungensis (Pope, 1934), Geoclemys kwangtungensis Pope, 1934, Emys
nigricans Gray, 1834, Clemmys nigricans (Gray, 1834), Clemmys schmackeri Boettger, 1894, Damonia
nigricans (Gray, 1834), Damonia mutica Boulenger, 1889
E: Red-necked Pond Turtle
China [3396, 5050]
Chinemys reevesii (Gray, 1831) III CN C EN
Synonyms: Geoclemys grangeri K. P. Schmidt, 1927, Geoclemys paracaretta Chang, 1929, Emys reevesii
Gray, 1831, Emys vulgaris picta Schlegel, 1844, Emys japonica Duméril, Bibron & Duméril, 1851, Damonia
unicolor Gray, 1873, Damonia reevesii (Gray, 1831)
E: Reeves's Turtle
CITES EC Reg. RL References
1
Canada (int) [3396]; China [3396, 5050]; Democratic People's Republic of Korea [3396]; Hong Kong, China
[5227]; Japan [3396]; Korea, Republic of [3396]; Taiwan, Province of China [3396, 5062]; United States (int)
[3396]
Chrysemys picta (Schneider, 1783) B
Synonym: Testudo picta Schneider, 1783
E: Painted Turtle
Canada [3396]; Mexico [3396]; United States [3396]
Clemmys insculpta (LeConte, 1830) II B VU 3396
Synonyms: Testudo insculpta LeConte, 1830, Emys speciosa Gray, 1831, Glyptemys insculpta (LeConte,
1830)
E: Wood Turtle, F: Clemmyde sculptée
Canada [3396]; United States [3396]
CITES Identification Manual Reference: A-301.007.007.002
Clemmys muhlenbergii (Schoepff, 1801) I A EN 3396
Synonyms: Testudo muhlenbergii Schoepff, 1801, Emys biguttata Say, 1824, Glyptemys muhlenbergii
(Schoepff, 1801), Clemmys nuchalis Dunn, 1917
E: Bog Turtle, Muhlenberg's Turtle, F: Clemmyde de Muhlenberg, Tortue de Muhlenberg, Tortue des
marécages, S: Galápago de Muhlenberg
United States [3396]
CITES Identification Manual Reference: A-301.007.007.004
Cuora amboinensis (Daudin, 1802) II B VU 4988, 5226
Synonyms: Terrapene bicolor Bell, 1825, Testudo amboinensis Daudin, 1802, Testudo melanocephala
Daudin, 1802, Emys cuoro Schweigger, 1812
E: Malaysian Box Turtle, South Asian Box Turtle, F: Tortue-boîte d'Asie orientale, S: Tortuga de caja malaya
Bangladesh [3396, 5228, 5236]; Brunei Darussalam; Cambodia [5143, 25563]; ?China [5050]; India [3396, 5175];
Indonesia [3396, 5094]: Bali [5094], Jawa [5094], Kalimantan [5094], Lesser Sunda Is [5094], Moluccas [5094],
Sulawesi [5094], Sumatera [5085, 5094]; Lao People's Democratic Republic [5037, 5052]; Malaysia [5095]:
Peninsular Malaysia [3396], Sabah [3396], Sarawak [3396]; Myanmar [3396, 5084, 10420]; Philippines [3396,
5098] ; Singapore; Thailand [3396, 5053, 6262]; Viet Nam [3396, 5173]
Cuora aurocapitata Luo & Zong, 1988 II B CR 4988
E: Golden-headed Box Turtle, Yellow-headed Box Turtle, F: Tortue-boîte à tête jaune
China [3396, 5050, 5106]
Cuora flavomarginata (Gray, 1863) II B EN 4988
Synonyms: Cistoclemmys flavomarginata Gray, 1863, Cyclemys flavomarginata (Gray, 1863)
E: Chinese Box Turtle, Yellow-margined Box Turtle, F: Tortue-boîte à bords jaune
China [3396, 5050, 5106, 10454]; Japan [3396]; Taiwan, Province of China [3396, 5062]: Taiwan [5062]
Cuora galbinifrons Bourret, 1939 II B CR 4988, 5226
Synonyms: Cuora hainanensis (Li, 1958), Cuora serrata Iverson & McCord, 1992, Cistoclemmys hainanensis
(Li, 1958), Cistoclemmys galbinifrons (Bourret, 1939), Cyclemys flavomarginata hainanensis Li, 1958,
Cyclemys hainanensis Li, 1958
E: Indochinese Box Turtle, F: Tortue-boîte à front jaune
China [3396, 5050]; Lao People's Democratic Republic [5037, 5052]; Viet Nam [3396, 5173]
Cuora mccordi Ernst, 1988 II B CR 4988
E: McCord's Box Turtle, F: Tortue-boîte de McCord
China [3396, 5050, 5106]
Cuora pani Song, 1984 II B CR 4988
Synonym: Cuora chriskarannarum Ernst & McCord, 1987
E: Pan's Box Turtle, F: Tortue-boîte de Pan
China [3396, 5050, 5106]
Cuora trifasciata (Bell, 1825) II B CR 4988, 5226
Synonyms: Sternotherus trifasciatus Bell, 1825, Cyclemys trifasciata (Bell, 1825)
E: Chinese Three-striped Box Turtle, F: Tortue-boîte à trois bandes
China [3396, 5050, 5106]; Hong Kong, China [5227]; ?Lao People's Democratic Republic [5037, 8469]; ?Myanmar
[3396]; Viet Nam [3396, 5173]
CITES EC Reg. RL References
2
Cuora yunnanensis (Boulenger, 1906) II B EX 4988, 25631
Synonym: Cyclemys yunnanensis Boulenger, 1906
E: Yunnan Box Turtle, F: Tortue-boîte du Yunnan
China [3396, 5050, 5106]: Yunnan
Cuora zhoui Zhao, Zhou & Ye, 1990 II B CR 4988
Synonym: Cuora pallidicephala McCord & Iverson, 1991
E: Zhou's Box Turtle, F: Tortue-boîte du Guangxi
China [3396, 5050, 5106]
Geoclemys hamiltonii (Gray, 1831) I A VU 3396
Synonyms: Melanochelys picta Murray, 1884, Emys hamiltonii Gray, 1831, Emys picquotii Lesson, 1831,
Emys guttata Gray, 1832, Damonia hamiltonii (Gray, 1831)
E: Black Pond Turtle, Black Spotted Turtle, Hamilton's Terrapin, Spotted Pond Turtle, F: émyde en toit,
Géoclemmyde d'Hamilton, Tortue à toit de l'Inde , Tortue de Hamilton, S: Galápago de Hamilton, Galápago
rayado
Bangladesh [3396, 5228]; India [3263, 3264, 3396, 5175, 7754]; ?Nepal [3396, 3526]; Pakistan [3396, 3521]
CITES Identification Manual Reference: A-301.007.013.001
Geoemyda spengleri (Gmelin, 1789) III CN C EN
Synonyms: Geoemyda japonica Fan, 1931, Testudo spengleri Gmelin, 1789, Testudo tricarinata Bory, 1804,
Nicoria spengleri (Gmelin, 1789)
E: Black-breasted Leaf Turtle
China [3396, 5050]; ?Indonesia [3396, 5094]: Sumatera; Japan [3396]; Viet Nam [3396, 5173]
Heosemys depressa (Anderson, 1875) II B CR
Synonyms: Geoemyda arakana Theobald, 1876, Geoemyda depressa (Anderson, 1875)
E: Arakan Forest Turtle, F: Héosémyde de l'Arakan
China; Myanmar [3396, 5084, 10420]
Heosemys grandis (Gray, 1860) II B VU
Synonym: Geoemyda grandis Gray, 1860
E: Giant Asian Pond Turtle, Orange-headed Temple Turtle, F: Héosémyde géante
Cambodia [3396, 5143, 25563]; Lao People's Democratic Republic [5037]; Malaysia [5095]: Peninsular Malaysia
[3396]; Myanmar [3396, 5084, 10420]; Thailand [3396, 5053, 6262]; Viet Nam [3396, 5173]
Heosemys leytensis Taylor, 1920 II B CR 25616
Synonym: Geoemyda leytensis (Taylor, 1920)
E: Leyte Pond Turtle, Philippine Pond Turtle, F: Héosémyde de Leyte
Philippines [3396, 5098, 13069, 25616]
Heosemys spinosa (Gray, 1831) II B EN
Synonyms: Geoemyda spinosa (Gray, 1831), Emys spinosa Bell, 1830
E: Spiny Terrapin, Spiny Turtle, Sunburst Turtle, F: Héosémyde épineuse
Brunei Darussalam [9338]; Indonesia [5094]: Kalimantan [3396], Sumatera [3396]; Malaysia [5095]: Peninsular
Malaysia [3396, 7361], Sabah [3396], Sarawak [3396]; Myanmar [5084, 10420]; Philippines; Thailand [3396, 5053]
Hieremys annandalii (Boulenger, 1903) II B EN
Synonym: Cyclemys annandalii Boulenger, 1903
E: Yellow-headed Temple Turtle
Cambodia [3396, 5143, 25563]; Lao People's Democratic Republic [5037]; Malaysia [5095]: Peninsular Malaysia;
Thailand [3396, 5053, 6262]; Viet Nam [3396, 5173]
Kachuga dhongoka (Gray, 1834) II B EN
Synonyms: Emys dhongoka Gray, 1834, Emys duvaucelii Duméril & Bibron, 1835, Kachuga hardwickii Gray,
1869
E: Dhongoka Roofed Turtle, Three-striped Roofed Turtle, F: Kachuga à cou rayé
Bangladesh [3396, 5228]; India [3396, 5175]; Nepal [3396]
Kachuga kachuga (Gray, 1831) II B CR
Synonyms: Batagur ellioti Gray, 1862, Emys kachuga Gray, 1831, Emys lineata Gray, 1831, Kachuga lineata
(Boulenger, 1889), Kachuga fusca Gray, 1870
E: Bengal Roof Turtle, Red-crowned Roofed Turtle, Sail Terrapin, F: Kachuga à front rouge
Bangladesh [3396, 5228]; India [3396, 5175]; Nepal [3396]
CITES EC Reg. RL References
3
Kachuga smithii (Gray, 1863) II B LR/nt
Synonym: Batagur smithii Gray, 1863
E: Brown River Turtle, Brown Roofed Turtle, F: Kachuga de Smith
Bangladesh [3396, 5228]; India [3396, 5175]; Nepal [3396]; Pakistan [3396]
Kachuga sylhetensis (Jerdon, 1870) II B EN
Synonyms: Pangshura sylhetensis Jerdon, 1870, Jerdonella sylhetensis (Jerdon, 1870)
E: Assam Roofed Turtle, Sylhet Roofed Turtle, F: Kachuga de l'Assam
Bangladesh [3396, 5228]; India [3396, 5175, 8733]
Kachuga tecta (Gray, 1831) I A LR/lc 3396
Synonyms: Emys tecta Gray, 1831, Emys trigibbosa Lesson, 1831, Pangshura dura Gray, 1869, Pangshura
flaviventer Günther, 1864, Pangshura ventricosa Gray, 1870
E: Dura Turtle, Indian Pond Turtle, Indian Roofed Turtle, Indian Sawback Turtle, Tent Turtle, F: Émyde
tricarénée, Kachuga à dos en toit, S: Galápago cubierto de la India, Tortuga india de visera
Bangladesh [3396, 5228, 5236]; India [3264, 3396, 5175, 7754]; Nepal [3266, 5133]; Pakistan [3396, 3521]
CITES Identification Manual Reference: A-301.007.019.005a
Kachuga tentoria (Gray, 1834) II B LR/lc
Synonyms: Emys tentoria Gray, 1834, Emys tecta intermedia Blanford, 1870, Kachuga intermedia
Boulenger, 1889, Pangshura leithii Gray, 1870, Pangshura tentoria (Gray, 1834)
E: Indian Tent Turtle, F: Kachuga carénée
Bangladesh [3396, 5228]; India [3396, 5175, 7754]; Nepal
Kachuga trivittata
(Duméril & Bibron, 1835) II B EN
Synonyms: Emys trivittata Duméril & Bibron, 1835, Kachuga peguensis Gray, 1869, Kachuga trilineata
Gray, 1869, Kachuga fusca Gray, 1870
E: Burmese Roofed Turtle
Myanmar [3396, 5084, 10420]
Leucocephalon yuwonoi
(McCord, Iverson & Boeadi, 1995) II B CR
Synonyms: Geoemyda yuwonoi McCord, Iverson & Boeadi, 1995, Heosemys yuwonoi (McCord, Iverson &
Boeadi, 1995)
E: Sulawesi Forest Turtle, F: Géoémyde des Célèbes
Indonesia: Sulawesi
Malayemys subtrijuga
(Schlegel & S. Müller, 1844) II B VU
Synonyms: Geoclemys macrocephala Gray, 1859, Emys subtrijuga Schlegel & S. Müller, 1844, Emys
nuchalis Blyth, 1863, Damonia crassiceps Gray, 1870, Damonia subtrijuga (Schlegel & S. Müller, 1844)
E: Malayan Snail-eating Turtle, Ricefield Turtle
Cambodia [5143, 25563]; China; Indonesia [5094]: Jawa [3396], Sumatera [3396]; Lao People's Democratic
Republic [5037]; Malaysia [5095]: Peninsular Malaysia [3396]; Thailand [3396, 5053, 6262]; Viet Nam [3396, 5173]
Mauremys iversoni Pritchard & McCord, 1991 III CN C DD 7634
E: Fujian Pond Turtle
China [3396, 5050]: Fujian, Guizhou
Mauremys mutica (Cantor, 1842) II B EN
Synonyms: Emys muticus Cantor, 1842, Clemmys mutica (Cantor, 1842)
E: Yellow Pond Turtle
China [3396, 5050]; Japan (int) [3396]; Taiwan, Province of China [3396, 5062]; Viet Nam [3396, 5173]
Mauremys pritchardi McCord, 1997 III CN C DD 6922
E: Pritchard's Pond Turtle
China [5050]; ?Myanmar [10420]
Melanochelys tricarinata (Blyth, 1856) I A VU 3396
Synonyms: Geoemyda tricarinata Blyth, 1856, Nicoria tricarinata (Blyth, 1856)
E: Three-keeled Land Tortoise, Three-keeled Land Turtle, Tricarinate Hill Turtle, F: Émyde indienne à trois
carènes, Émyde ocellée de Birmanie, Géoémyde tricarénée, Tortue tricarénée, S: Galápago terrestre,
Galápago tricarenado
CITES EC Reg. RL References
4
Bangladesh [3396, 5228, 5236]; India [3264, 3396, 5175]; ?Nepal [3526, 5133]
CITES Identification Manual Reference: A-301.007.023.001
Melanochelys trijuga (Schweigger, 1812) D LR/nt
Synonyms: Geoemyda trijuga (Schweigger, 1812), Emys trijuga Schweigger, 1812
E: Indian Black Turtle
Bangladesh [3396, 5228]; British Indian Ocean Territory (int?) [3396]; India [3396, 5175]; Maldives (int?) [3396];
Myanmar [3396, 5084, 7414, 10420]; Nepal [3396]; Sri Lanka [3396]
Morenia ocellata (Duméril & Bibron, 1835) I A VU 3396, 5226
Synonyms: Emys ocellata Duméril & Bibron, 1835, Emys berdmorei Blyth, 1859
E: Bengal Eyed Terrapin, Burmese Eyed Turtle, Burmese Swamp Turtle, Peacock Turtle, Swamp Turtle,
F: Émyde à ocelles de Birmanie, Terrapenes , Tortue de Birmanie, S: Galápago de Birmania, Galápago
ocelado de Birmania
Myanmar [3396, 5084, 7414, 10420]
CITES Identification Manual Reference: A-301.007.024.001
Notochelys platynota (Gray, 1834) II B VU
Synonyms: Emys platynota Gray, 1834, Cyclemys giebelii Hubrecht, 1881, Cyclemys platynota (Gray, 1834)
E: Malayan Flat-shelled Turtle
Brunei Darussalam [25611]; Indonesia [5094]: Jawa [3396], Kalimantan [3396], Sumatera [3396]; Malaysia
[5095]: Peninsular Malaysia [3396], Sabah [3396], Sarawak [3396]; ?Myanmar [3396]; Singapore; Thailand
[3396, 5053]; Viet Nam [3396]
Ocadia glyphistoma McCord & Iverson, 1994 III CN C DD 7366
E: Notch-mouthed Stripe-necked Turtle
?China [5050]; ?Viet Nam
Ocadia philippeni McCord & Iverson, 1992 III CN C DD 7367
E: Philippen's Stripe-necked Turtle
China [3396, 5050]: Hainan
Ocadia sinensis (Gray, 1834) III CN C EN
Synonyms: Emys sinensis Gray, 1834, Emys bennettii Gray, 1844
E: Chinese Stripe-necked Turtle
China [3396, 5050]; ?Lao People's Democratic Republic [3396]; Taiwan, Province of China [3396, 5062]; Viet
Nam [3396, 5173]
Orlitia borneensis Gray, 1873 II B EN
Synonyms: Hardella baileyi Bartlett, 1895, Bellia borneensis (Gray, 1873), Adelochelys crassa Baur, 1896,
Liemys inornata Boulenger, 1897, Clemmys gibbera Peters, 1874
E: Bornean River Turtle, Malaysian Giant Terrapin, Malaysian Giant Turtle, F: Emyde géante de Borneo
Indonesia [5094]: Kalimantan [3396], Sumatera [3396]; Malaysia [5095]: Peninsular Malaysia [3396, 7361],
Sarawak [3396]
Pyxidea mouhotii (Gray, 1862) II B EN
Synonym: Cyclemys mouhotii Gray, 1862
E: Jagged-shelled Turtle, Keel-backed Turtle, Keeled Box Turtle
China [3396, 5050]; India [3396, 5175]; Lao People's Democratic Republic [5037]; Myanmar [3396, 5084, 10420];
?Thailand [3396, 5053]; Viet Nam [3396, 5173]
Sacalia bealei (Gray, 1831) III CN C EN
Synonyms: Cistuda bealei Gray, 1831, Clemmys bealei (Gray, 1831), Mauremys bealei (Gray, 1831)
E: Beal's Eyed Turtle
China [3396, 5050]; Hong Kong, China (int?) [3396, 5227]
Sacalia pseudocellata
Iverson & McCord, 1992 III CN C DD 7368
E: Chinese False-eyed Turtle
China [5050]
Sacalia quadriocellata (Siebenrock, 1903) III CN C EN
Synonym: Clemmys bealei quadriocellata Siebenrock, 1903
E: Four-eyed Turtle
China [3396, 5050]; Lao People's Democratic Republic [5037]; Viet Nam [3396, 5173]
CITES EC Reg. RL References
5
China [3396, 5050]; Lao People's Democratic Republic [5037]; Viet Nam [3396, 5173]
Siebenrockiella crassicollis (Gray, 1831) II B VU
Synonyms: Emys nigra Blyth, 1855, Emys crassicollis Gray, 1831, Pangshura cochinchinensis Tirant, 1885,
Bellia crassicollis Boulenger, 1889
E: Black Marsh Turtle
Cambodia [5143, 25563]; Indonesia [5094]: Jawa [3396], Kalimantan [3396], Sumatera [3396]; ?Lao People's
Democratic Republic [5037]; Malaysia [5095]: Peninsular Malaysia [3396]; Myanmar [3396, 5084, 10420];
Thailand [3396, 5053]; Viet Nam [3396, 5173]
Terrapene carolina (Linnaeus, 1758) II B LR/nt 3396
Synonyms: Terrapene mexicana (Gray, 1849), Terrapene goldmani Stejneger, 1933, Terrapene yucatana
(Boulenger, 1895), Terrapene maculata Bell, 1825, Terrapene nebulosa Bell, 1825, Terrapene bauri Taylor,
1895, Testudo virgulata Latreille, 1801, Testudo incarcerata Bonnaterre, 1789, Testudo incarceratostriata
Bonnaterre, 1789, Testudo clausa Gmelin, 1789, Testudo carolina Linnaeus, 1758, Testudo carinata
Linnaeus, 1758, Testudo brevicaudata Lacépède, 1788, Monoclida kentukensis Rafinesque, 1822, Cistudo
triunguis Agassiz, 1857, Cistudo mexicana Gray, 1849, Cistudo yucatana Boulenger, 1895, Cistudo carolina
(Linnaeus, 1758), Cistudo virginea Agassiz, 1857, Cistudo major Agassiz, 1857, Emys schneideri
Schweigger, 1812, Emys kinosternoides Gray, 1831
E: Common Box Turtle, Eastern Box Turtle, F: Tortue-boîte de Caroline
Canada [3396]; Mexico [3395, 3396]; United States [3163, 3395, 3396]
Terrapene coahuila
K. P. Schmidt & Owens, 1944 I A EN 3396
E: Aquatic Box Turtle, Coahuila Box Turtle, Coahuila Turtle, Water Box Turtle, F: Tortue-boîte de Coahuila,
Tortue-boîte du Mexique, S: Galápago acuático encajado, Galápago caja mexicano, Tortuga Coahuila
Mexico [3396]
(Cuatro Cienegas basin)
CITES Identification Manual Reference: A-301.007.031.002
Terrapene nelsoni Stejneger, 1925 II B DD 3396
Synonym: Terrapene klauberi Bogert, 1943
E: Spotted Box Turtle, F: Tortue-boîte du Mexique
Mexico [3396]
Terrapene ornata (Agassiz, 1857) II B LR/nt 3396
Synonym: Cistudo ornata Agassiz, 1857
E: Ornate Box Turtle, Western Box Turtle, F: Tortue-boîte ornée commune
Mexico [3395, 3396]; United States [3163, 3395, 3396]
Trachemys scripta (Schoepff, 1792) B/NR LR/nt
Synonyms: Testudo scripta Schoepff, 1792, Testudo serrata Daudin, 1802, Chrysemys scripta (Schoepff,
1792), Pseudemys scripta (Schoepff, 1792)
E: Common Slider
Bahrain (int) [9832]; Belize [12317]; Colombia [3396]; Costa Rica [3396, 7830]; Cyprus (int); El Salvador;
France (int) [3396]; Germany (int) [3396]; Guadeloupe (int) [3396]; Guam (int); Guatemala [3396]; Honduras
[3396, 25560]; Hong Kong, China (int) [5227]; Indonesia (int) [5094]: Irian Jaya (int), Jawa (int), Kalimantan
(int), Sulawesi (int), Sumatera (int); Israel (int) [3396]; Italy (int); Japan (int); Mexico [3396]; Myanmar
(int) [10420]; Netherlands (int) [3396]; Nicaragua [3396]; Northern Mariana Islands (int); Panama [3396];
Singapore (int); South Africa (int) [3396]; Spain (int) [25662]; Taiwan, Province of China (int) [5062];
Thailand (int) [5053]; United States [3396]; Venezuela [3396]
Non-EC Reg. populations LR/nt
Belize; Colombia; Costa Rica; El Salvador; Guadeloupe (int); Guatemala; Honduras; Mexico; Nicaragua;
Panama; United States; Venezuela
Trachemys scripta elegans
(Wied, 1839) B
Synonyms: Trachemys lineata Gray, 1873, Chrysemys scripta elegans (Wied, 1839), Emys elegans Wied,
1839, Emys holbrookii Gray, 1844, Emys sanguinolenta Gray, 1855
E: Red-eared Slider, Red-eared terrapin, F: Trachémyde à tempes rouges
Bahrain (int); Cyprus (int); France (int); Germany (int); Guam (int); Hong Kong, China (int); Indonesia
(int); Israel (int); Italy (int); Japan (int); Mexico; Myanmar (int) [10420]; Netherlands (int); Northern
Mariana Islands (int); Singapore (int); South Africa (int); Spain (int) [25662]; Taiwan, Province of China
(int); Thailand (int); United States
CITES EC Reg. RL References
6
Family: TESTUDINIDAE
Chersina angulata (Schweigger, 1812) II B 3396
Synonyms: Testudo tabulata africana Schweigger, 1812, Testudo bellii Gray, 1828, Testudo angulata
Schweigger, 1812
E: Angulated Tortoise, Bowsprit Tortoise, South African Bowsprit Tortoise, F: Tortue à soc d'Afrique du Sud,
S: Tortuga ariete, Tortuga de pechera
Namibia [3396]; South Africa [3197, 3396, 3674]
CITES Identification Manual Reference: A-301.011.002.001
Geochelone carbonaria (Spix, 1824) II B 3396, 3582, 3639
Synonyms: Testudo carbonaria Spix, 1824, Testudo boiei Wagler, 1833, Chelonoides carbonaria (Spix, 1824)
E: Red-footed Tortoise, Wood Tortoise, F: Tortue charbonnière, S: Jabuti, Karumbé, Tortuga de patas rojas
Anguilla (int); Antigua and Barbuda (int); Argentina [3396]; Barbados (ex, int) [3254]; Bolivia [3396, 5116,
25612]; Brazil [3396]; British Virgin Islands (int); Colombia [3396]; Dominica (int) [3254]; French Guiana
[3396]; Grenada (ex?, int) [11881]; Guyana [3396, 25473, 25581]; Montserrat (int); Panama [3396]; Paraguay
[3396, 10110]; Saint Vincent and the Grenadines (int); Suriname [3396]; Trinidad and Tobago; United States
Virgin Islands (int); Venezuela [3396, 3582, 5115]
CITES Identification Manual Reference: A-301.011.003.001
Geochelone chilensis (Gray, 1870) II B VU 3396, 3464, 3711
Synonyms: Geochelone donosobarrosi Freiberg, 1973, Geochelone petersi Freiberg, 1973, Testudo chilensis
Gray, 1870, Testudo argentina Sclater, 1870, Chelonoides chilensis (Gray, 1870)
E: Argentine Tortoise, Chaco Tortoise, Southern Wood Tortoise, F: Tortue de la pampa, S: Tortuga terrestre
argentina
Argentina [3396]; ?Bolivia [12403]; Paraguay [3396, 10110]
CITES Identification Manual Reference: A-301.011.003.002
Geochelone denticulata (Linnaeus, 1766) II B VU 3396, 3582
Synonyms: Testudo hercules Spix, 1824, Testudo sculpta Spix, 1824, Testudo cagado Spix, 1824, Testudo
denticulata Linnaeus, 1766, Testudo tabulata Schoepff, 1792, Testudo tessellata Schneider, 1792,
Chelonoides denticulata (Linnaeus, 1766)
E: Brazilian Giant Tortoise, Forest Tortoise, South American Tortoise, South American Yellow-footed
Tortoise, Yellow-footed Tortoise, F: Tortue de l'Amérique du sud, Tortue dentelée, Tortue denticulée,
S: Motelo, Tortuga de patas amarillas
Bolivia (int) [3396, 25612]; Brazil [3396]; Colombia [3396]; Dominica (int) [3254]; Ecuador [3396, 3523]; French
Guiana [3396]; Guadeloupe [12496]: Guadeloupe; Guyana [3396, 25473, 25581]; Peru [3281, 3396, 3601, 12589];
Suriname [3396]; Trinidad and Tobago [3396]; Venezuela [3396, 3582, 5115]
CITES Identification Manual Reference: A-301.011.003.003
Geochelone elegans (Schoepff, 1795) II B LR/lc 3396
Synonyms: Testudo megalopus Blyth, 1853, Testudo elegans Schoepff, 1795, Testudo stellata Schweigger,
1812, Testudo actinoides Bell, 1828, Chelonoides elegans (Schoepff, 1795)
E: Indian Star Tortoise, Star Tortoise, F: Tortue étoilée de l'Inde, S: Tortuga estrellada de la India
India [3263, 3264, 3396, 5175]; ?Myanmar [10420]; Pakistan [3396, 3521]; Sri Lanka [3396]
CITES Identification Manual Reference: A-301.011.003.004
Geochelone gigantea (Schweigger, 1812) II B VU 3396
Synonyms: Testudo elephantina Duméril & Bibron, 1835, Testudo gigantea Schweigger, 1812, Testudo
hololissa Günther, 1877, Testudo daudinii Duméril & Bibron, 1835, Aldabrachelys elephantina (Duméril &
Bibron, 1835), Chelonoides gigantea (Schweigger, 1812), Dipsochelys elephantina (Duméril & Bibron,
1835), Dipsochelys daudinii (Duméril & Bibron, 1835), Dipsochelys dussumieri (Gray, 1831), Dipsochelys
grandidieri (Vaillant, 1885), Dipsochelys arnoldi Bour, 1982, Dipsochelys hololissa (Günther, 1877),
Dipsochelys abrupta (Grandidier, 1868)
E: Aldabra Giant Tortoise, F: Tortue géante, Tortue géante d'Aldabra, S: Tortuga gigante de Aldabra
Mauritius (int) [3396]; Réunion (int) [3396]; Seychelles: Seychelles (int), Aldabra [3396]; United Republic of
Tanzania (int) [8517]
CITES Identification Manual Reference: A-301.011.003.009
Geochelone nigra (Quoy & Gaimard, 1824) I A VU 3396, 3596
Synonyms: Geochelone elephantopus (Harlan, 1827), Testudo macrophyes Garman, 1917, Testudo
vandenburghi De Sola, 1930, Testudo elephantopus Harlan, 1827, Testudo typica Garman, 1917, Testudo
microphyes Günther, 1875, Testudo vicina Günther, 1875, Testudo nigra Quoy & Gaimard, 1824, Testudo
planiceps Gray, 1854, Testudo clivosa Garman, 1917, Chelonoides nigra (Quoy & Gaimard, 1824)
CITES EC Reg. RL References
7
E: Galapagos Giant Tortoise, F: Tortue éléphantine, Tortue géante des Galapagos, S: Tortuga de las
Galápagos, Tortuga gigante de las Galápagos
Ecuador [3396]: Galapagos [5198]
CITES Identification Manual Reference: A-301.011.003.005
Geochelone pardalis (Bell, 1828) II B 3464, 4988, 8735
Synonym: Testudo pardalis Bell, 1828
E: Leopard Tortoise, Mountain Tortoise, F: Tortue-léopard du Cap, S: Tortuga leopardo
Angola [3396]; Botswana [3146, 3396]; Democratic Republic of the Congo; Djibouti [5135]; Ethiopia [3396];
Kenya [3396, 8517]; Malawi [3396]; Mozambique [3396]; Namibia [3396]; Somalia [3396, 5135]; South Africa
[3396, 5159]; Sudan [3396]; ?Swaziland [5038]; Uganda [3396, 8517]; United Republic of Tanzania [3396, 5136,
7273, 8517, 25472] ; Zambia [3396, 5206]; Zimbabwe [3396, 5100]
CITES Identification Manual Reference: A-301.011.003.011
Geochelone platynota (Blyth, 1863) II B CR 3396, 5226
Synonym: Testudo platynota Blyth, 1863
E: Burmese Starred Tortoise, Flatback Tortoise, F: Tortue étoilée de Birmanie, S: Tortuga estrellada de
Burma
Myanmar [3396, 5084, 8466, 10420]
CITES Identification Manual Reference: A-301.011.003.012
Geochelone radiata (Shaw, 1802) I A VU 3396
Synonyms: Testudo radiata Shaw, 1802, Testudo hypselonota Bourret, 1941 , Testudo coui Daudin, 1802,
Testudo desertorum Grandidier, 1869, Asterochelys radiata (Shaw, 1802)
E: Radiated Tortoise, F: Tortue radiée de Madagascar, Tortue rayonnée, S: Tortuga estrellada de
Madagascar, Tortuga rayada
Madagascar [3396, 4995, 6516]; Mauritius (int) [3396]; Réunion (int) [3396]
CITES Identification Manual Reference: A-301.011.003.013
Geochelone sulcata (Miller, 1779) II B VU 3396
Synonyms: Testudo sulcata Miller, 1779, Testudo calcarata Schneider, 1784, Testudo radiata senegalensis
Gray, 1831
E: African Spurred Tortoise, Grooved Tortoise, F: Tortue sillonnée, S: Tortuga con púas
Chad [3396, 12975]; Egypt; Eritrea [79, 3396]; Ethiopia [3396]; Mali [3396, 5040, 8450]; Mauritania [3396]; Niger
[12588] ; Nigeria [3396]; Senegal [3396, 8734]; ?Somalia [3396]; Sudan [3396]
CITES Identification Manual Reference: A-301.011.003.014
Geochelone yniphora (Vaillant, 1885) I A EN 3262, 3396
Synonyms: Testudo yniphora Vaillant, 1885, Asterochelys yniphora (Vaillant, 1885)
E: Angonoka, Madagascar Angulated Tortoise, Madagascar Tortoise, F: Tortue à éperon, Tortue à plastron
éperonné, Tortue à soc, Tortue à soc de Madagascar, Tortue de Madagascar, S: Tortuga de espolones,
Tortuga de Madagascar, Tortuga globulosa malagache
Madagascar [3396, 4995]
CITES Identification Manual Reference: A-301.011.003.016
Gopherus agassizii (Cooper, 1863) II B VU 3141, 3191, 3396, 3564
Synonyms: Testudo agassizii (Cooper, 1863), Gopherus polyphemus agassizii (Cooper, 1863), Scaptochelys
agassizii (Cooper, 1863), Xerobates agassizii (Cooper, 1863)
E: Desert Tortoise, F: Gophère d'Agassiz, Tortue d'Agassiz, S: Tortuga del desierto
Mexico [3396, 3557]; United States [3396, 3460]
CITES Identification Manual Reference: A-301.011.004.001
Gopherus berlandieri (Agassiz, 1857) II B LR/lc 3191, 3396, 3608
Synonyms: Testudo berlandieri (Agassiz, 1857), Gopherus polyphemus berlandieri (Agassiz, 1857),
Scaptochelys berlandieri (Agassiz, 1857), Xerobates berlandieri (Agassiz, 1857)
E: Berlandier's Tortoise, Texas Tortoise, F: Gophère du Texas, Tortue du Texas, S: Tortuga de Texas
Mexico [3396]; United States [3396]
CITES Identification Manual Reference: A-301.011.004.002
Gopherus flavomarginatus Legler, 1959 I A VU 3396, 3530
Synonym: Gopherus polyphemus flavomarginatus Legler, 1959
CITES EC Reg. RL References
8
E: Bolson Tortoise, Mexican Giant Tortoise, Yellow-bordered Tortoise, F: Gophère , Gophère à bords jaunes,
Tortue à bords jaunes, Tortue du Mexique, Tortue fouisseuse du Mexique, S: Tortuga grande, Tortuga
llanera, Tortuga topo
Mexico [3396]
CITES Identification Manual Reference: A-301.011.004.003
Gopherus polyphemus (Daudin, 1802) II B VU 3152, 3396
Synonyms: Testudo polyphemus Daudin, 1802, Testudo depressa Guérin-Méneville, 1829, Testudo gopher
Gray, 1844
E: Gopher Tortoise, F: Gophère polyphème, Tortue de la Floride, Tortue gaufrée, S: Tortuga terrestre de
Florida
United States [3396]
CITES Identification Manual Reference: A-301.011.004.004
Homopus areolatus (Thunberg, 1787) II B 3189, 3354, 3396
Synonyms: Testudo areolata Thunberg, 1787, Testudo miniata Lacépède, 1788, Testudo fasciata Daudin,
1802, Chersine tetradactyla Merrem, 1820
E: Areolated Tortoise, Beaked Cape Tortoise, Grooved Mountain Tortoise, Parrot-beaked Tortoise,
F: Homopode aréolé, S: Tortuga pico de loro
South Africa [3396]
CITES Identification Manual Reference: A-301.011.005.001
Homopus boulengeri Duerden, 1906 II B 3189, 3396
E: Boulenger's Cape Tortoise, Boulenger's Padloper, Karoo Padloper, F: Homopode de Boulenger,
S: Tortuga terrestre
South Africa [3396]
CITES Identification Manual Reference: A-301.011.005.002
Homopus femoralis Boulenger, 1888 II B 3189, 3396
E: Greater Padloper, Karoo Cape Tortoise, F: Homopode à éperon, Homopode éperonné
Lesotho; South Africa [3396]
CITES Identification Manual Reference: A-301.011.005.003
Homopus signatus (Schoepff, 1801) II B LR/nt 3189, 3396
Synonyms: Testudo signata Schoepff, 1801, Testudo cafra Daudin, 1802, Testudo juvencella Daudin, 1831,
Pseudomopus signatus (Schoepff, 1801)
E: Speckled Cape Tortoise, Speckled Padloper, Speckled Tortoise, F: Homopode marqué, S: Tortuga
manchada
?Namibia; South Africa [3396]
CITES Identification Manual Reference: A-301.011.005.004
Indotestudo elongata (Blyth, 1853) II B EN 3396, 5226
Synonyms: Geochelone elongata (Blyth, 1853), Testudo elongata Blyth, 1853, Testudo parallelus
Annandale, 1913
E: Elongated Tortoise, Pineapple Tortoise, Red-nosed Tortoise, Yellow Tortoise, Yellow-headed Tortoise,
F: Tortue à tête jaune
Bangladesh [3396, 5228, 5236]; Cambodia [3396, 5143, 25563]; China [3396, 5050, 5106]: Guangxi [3574]; India
[3264, 3396, 5175] ; Lao People's Democratic Republic [3396, 5037, 5052]; Malaysia [3186, 5095]: Peninsular
Malaysia [3396]; Myanmar [3396, 5084, 10420]; Nepal [3396, 3678, 5133]; Thailand [3396, 5053, 6262]; Viet Nam
[3396, 5173]
CITES Identification Manual Reference: A-301.011.003.006
Indotestudo forstenii
(Schlegel & S. Müller, 1844) II B EN 3381, 3396, 5226
Synonyms: Geochelone travancorica (Boulenger, 1907), Geochelone forstenii (Schlegel & S. Müller, 1844),
Testudo travancorica Boulenger, 1907, Testudo forstenii Schlegel & S. Müller, 1844, Indotestudo
travancorica (Boulenger, 1907)
E: Celebes Tortoise, Forsten's Tortoise, Travancore Tortoise, F: Tortue de Tranvancore, Tortue des Célèbes,
S: Tortuga marrón de la India
India [3264, 3396]; Indonesia [3396]: Moluccas [3396], Sulawesi [3396, 5094, 8465]
CITES Identification Manual Reference: A-301.011.003.008
Kinixys belliana Gray, 1831 II B 3396
CITES EC Reg. RL References
9
Synonyms: Homopus darlingi Boulenger, 1902, Homopus nogueyi Lataste, 1886, Cinixys belliana (Gray,
1831), Cinixys lobatsiana Power, 1927, Cinixys dorri Lataste, 1888, Kinixys youngi Hewitt, 1931, Kinixys
lobatsiana (Power, 1927), Kinixys spekii Gray, 1863, Kinixys schoensis Rüppell, 1845, Kinixys australis
Hewitt, 1931, Kinixys jordani Hewitt, 1931
E: Bell's Hinged Tortoise, Bell's Hinged-backed Tortoise, F: Kinixys de Bell
Angola [3396]; Benin [3396]; Botswana [3146, 3396]; Burkina Faso [3396, 6235]; ?Burundi [6235, 25523];
Cameroon [3396, 25340, 25380]; Central African Republic [3396, 6235]; Congo; Côte d'Ivoire; Democratic
Republic of the Congo [3396, 6235]; ?Djibouti; Eritrea [79, 3396]; Ethiopia [3396, 6235]; Gabon; Gambia [3363,
3396] ; Ghana [3396]; Guinea [3396]; Guinea-Bissau [3396]; Kenya [3396, 8517]; ?Liberia; Madagascar (int)
[3396, 3431]; Malawi [3396, 6235]; Mali [3396, 5040]; ?Mauritania; Mozambique [3396, 6235]; Namibia; Niger
[12588] ; Nigeria [3396]; Rwanda [3374, 8517]; Senegal [3396]; Sierra Leone [3396]; Somalia [3396, 6235]; South
Africa [3396]: Natal [6235]; Sudan [3396]; Swaziland [3396, 6235]; Togo [3396]; Uganda [3396, 8517]; United
Republic of Tanzania [3396, 8517]; Zambia [3396, 6235]; Zimbabwe [3205, 3396]
CITES Identification Manual Reference: A-301.011.006.001
Kinixys erosa (Schweigger, 1812) II B DD 3396
Synonyms: Testudo erosa Schweigger, 1812, Cinixys erosa (Schweigger, 1812), Kinixys castanea Bell, 1827
E: Common Tortoise, Forest Hinged Tortoise, Schweigger's Tortoise, Serrated Hinge-back Tortoise, Serrated
Hinge-backed Tortoise, Serrated Tortoise, F: Kinixys rongée, Tortue articulée d'Afrique
Angola [3396, 6235]; Benin [3396]; Burkina Faso [6235]; Cameroon [3396, 5120, 25340, 25354, 25379]; Central
African Republic [6235]; Congo [3396, 8749]; Côte d'Ivoire [3396, 6235, 12544]; Democratic Republic of the Congo
[3396, 6235]; Equatorial Guinea [3396, 6235, 25443]; Gabon [3396, 6235, 10390, 10393]; Gambia [3363, 3396]; Ghana
[3396, 6235]; Guinea [6235]; ?Guinea-Bissau; Liberia [3396, 6235]; Nigeria [3396, 6235]; ?Rwanda [6235, 8517];
Senegal; Sierra Leone [3396, 6235]; ?Togo [6235]; Uganda [3396, 8517]
CITES Identification Manual Reference: A-301.011.006.002
Kinixys homeana Bell, 1827 II B DD 3396
Synonym: Cinixys homeana (Bell, 1827)
E: Home's Hinge-back Tortoise, Home's Hinged Tortoise, Home's Hinged-backed Tortoise, F: Kinixys de
Home
?Benin [6235]; Cameroon [3396, 5120, 25340, 25354, 25379]; Congo; Côte d'Ivoire [3396, 6235]; Democratic
Republic of the Congo [3396, 6235]; Equatorial Guinea [3396, 6235]; ?Gabon [6235]; Ghana [3396, 6235]; Liberia
[3396, 6235]; Nigeria [3396, 6235]; Sierra Leone [12527]; ?Togo [6235]
CITES Identification Manual Reference: A-301.011.006.003
Kinixys natalensis Hewitt, 1935 II B LR/nt 3396
Synonym: Kinixys belliana natalensis Hewitt, 1935
E: Natal Hinge-back Tortoise, Natal Hinge-backed Tortoise, Natal Hinged Tortoise, F: Kinixys du Natal
Mozambique [3396, 6235]; South Africa [3396, 6235]; Swaziland [3396, 6235]
Malacochersus tornieri (Siebenrock, 1903) II A VU 3396, 3464
Synonyms: Testudo tornieri Siebenrock, 1903, Testudo loveridgii Boulenger, 1920, Testudo procterae
Loveridge, 1923
E: Crevice Tortoise, Pancake Tortoise, Softshell Tortoise, Tornier's Tortoise, F: Tortue à carapace souple,
Tortue de Tornier, S: Tortuga de cuña
Kenya [3396, 6235, 8517]; United Republic of Tanzania [3396, 5039, 5073, 8517]; ?Zimbabwe (int)
CITES Identification Manual Reference: A-301.011.007.001
Manouria emys (Schlegel & S. Müller, 1844) II B EN 3396, 5226, 12373
Synonyms: Geochelone emys (Schlegel & S. Müller, 1844), Testudo falconeri Gray, 1869, Testudo nutapundi
(Reimann in Wirot, 1979), Testudo emys Schlegel & S. Müller, 1844, Testudo emydoides Duméril, Bibron &
Duméril, 1851, Testudo phayrei Blyth, 1853, Teleopus luxatus LeConte, 1854, Manouria fusca Gray, 1854
E: Asian Giant Tortoise, Asian Tortoise, Black Giant Tortoise, Burmese Brown Tortoise, Burmese Mountain
Tortoise, Six-legged Tortoise, F: Tortue brune
Bangladesh [3396, 5228, 5236, 6235]; ?Cambodia [12587]; India [3264, 3396, 5175]: Assam [6235]; Indonesia
[6235]: Kalimantan [3396, 5094], Sumatera [3396, 5085, 5094]; Malaysia [5095]: Peninsular Malaysia [3186, 3396,
7361] , Sabah [3396, 8085], Sarawak [3396]; Myanmar [3396, 5084, 7414, 10420]; Singapore [10417]; Thailand [3396,
5053, 5081, 6235, 6262] ; Viet Nam [3396]
CITES Identification Manual Reference: A-301.011.003.007
Manouria impressa (Günther, 1882) II B VU 3396, 5226
Synonyms: Geochelone impressa (Günther, 1882), Geoemyda impressa Günther, 1882, Geoemyda
latinuchalis Vaillant, 1894, Testudo impressa (Günther, 1882), Testudo pseudemys Boulenger, 1903
E: Impressed Tortoise, F: Tortue imprimée, S: Tortuga marrón de Burma
CITES EC Reg. RL References
10
Cambodia [5143, 25563]; ?China [5050, 5106]: Hunan [6235]; Lao People's Democratic Republic [5037, 5052];
Malaysia [5095]: Peninsular Malaysia [3396]; Myanmar [3396, 10420, 10431]; Thailand [3396, 5053, 5132, 6262];
Viet Nam [3396, 5173]
CITES Identification Manual Reference: A-301.011.003.010
Psammobates geometricus (Linnaeus, 1758) I A EN 3354, 3396
Synonyms: Testudo luteola Daudin, 1802, Testudo strauchi Lidth de Jeude, 1893, Testudo geometrica
Linnaeus, 1758
E: Geometric Tortoise, F: Sakafi, Tortue géométrique, S: Sacafi, Tortuga geométrica
South Africa [3396]: Cape Province [6235]
CITES Identification Manual Reference: A-301.011.008.001
Psammobates oculiferus (Kuhl, 1820) II B 3396
Synonyms: Testudo oculifera Kuhl, 1820, Testudo semiserrata A, Smith, 1839
E: African Serrated Tortoise, Kuhl's Tortoise, Serrated Star Tortoise, Serrated Tent Tortoise, Toothed Cape
Tortoise, F: Tortue ocelée, S: Tortuga estrellada sudafricana de caparazón
Botswana [3146, 3396]; Namibia [3396]; South Africa [3396, 6235]: Orange Free state, Transvaal
CITES Identification Manual Reference: A-301.011.008.002
Psammobates tentorius (Bell, 1828) II B 3396
Synonyms: Homopus bergeri Lindholm, 1906, Testudo tentoria Bell, 1828, Testudo boettgeri Siebenrock,
1904, Testudo oscarboettgeri Lindholm, 1929, Testudo geometrica nigriventris Gray, 1855, Testudo smithi
Boulenger, 1886, Testudo verreauxii (Gray, 1870), Testudo seimundi Boulenger, 1903, Testudo trimeni
Boulenger, 1886, Testudo verroxii A. Smith, 1839, Testudo fiski Boulenger, 1886, Chersinella tentoria (Bell,
1828), Chersinella fiski (Boulenger, 1886), Chersinella schonlandi Hewitt, 1934, Chersinella verroxii (A.
Smith, 1839), Peltastes verreauxii Gray, 1870, Psammobates depressa FitzSimons, 1938
E: African Tent Tortoise, South African Tent Tortoise, F: Tortue bosselée, S: Tortuga estrellada sudafricana
Namibia [3396]; South Africa [3354, 3396]
CITES Identification Manual Reference: A-301.011.008.003
Pyxis arachnoides Bell, 1827 I A VU 3396
Synonym: Pyxis madagascariensis Lesson, 1831
E: Spider Tortoise, F: Pyxide arachnoide, S: Tortuga araña, Tortuga de plastrón articulado
Madagascar [3396, 6516]
CITES Identification Manual Reference: A-301.011.009.001
Pyxis planicauda (Grandidier, 1867) I A EN 3396
Synonyms: Testudo planicauda Grandidier, 1867, Acinixys planicauda Grandidier, 1867
E: Flat-backed Spider Tortoise, Flat-shelled Spider Tortoise, Madagascar Flat-shelled Tortoise, Malagasy
Flat-tailed Tortoise, F: Pyxide à dos plat, Pyxide à queue platte, S: Tortue de cola plana
Madagascar [3396, 6235]
CITES Identification Manual Reference: A-301.011.001.001
Testudo graeca Linnaeus, 1758 II A VU 3396, 3436, 3464, 5137
Synonyms: Testudo nabeulensis (Highfield, 1990), Testudo whitei Bennett in White, 1836, Testudo
flavominimaralis Highfield & Martin, 1990, Testudo mauritanica Duméril & Bibron, 1835, Testudo zarudnyi
Nikolsky, 1896, Testudo antakyensis Perälä, 1996, Testudo perses Perälä, 2002, Testudo soussensis Pieh,
2001, Testudo pusilla Linnaeus, 1758, Testudo anamurensis Weissinger, 1987, Testudo armeniaca
Chkhikvadze & Bakradze, 1991, Testudo cyrenaica Pieh & Perälä, 2002, Furculachelys nabeulensis Highfield,
1990, Furculachelys whitei (Bennett, 1836)
E: Algerian Tortoise, Common Tortoise, Greek Tortoise, Moorish Tortoise, Spur-thighed Tortoise, F: Tortue
grecque , Tortue grecque, Tortue mauresque, Tortue mauresque, S: Tortuga mora, Tortuga mora
Albania [5161]; Algeria [3396, 8277, 10377]; Armenia [3396, 8437]; Azerbaijan [3396, 8437]; Bulgaria [3167, 3396];
Cyprus (int) [3377]; Egypt [3396]; France (int) [3396]; Georgia [3396]; Greece [3396]: East Aegean Is; Iran
(Islamic Republic of) [3130, 3396, 8437, 9832]; Iraq [3396, 3470, 9832]; Israel [3396, 3500]; Italy (int) [3396]:
Sardegna, Sicilia (int); Jordan; ?Lebanon; Libyan Arab Jamahiriya [3396, 3632]; Moldova, Republic of [3396];
Morocco [3396, 3435, 5131, 10377]; Romania [3321, 3396]; Russian Federation [3396, 5074]; Serbia and
Montenegro; Spain [3396]: Spain [3396], Canary Is (int) [3396, 6235]; Syrian Arab Republic [3396]; The former
Yugoslav Republic of Macedonia [3396]; Tunisia [3396, 10377]; Turkey [3161, 3396, 8169, 8437, 9832, 25184, 25185];
Turkmenistan [3396]; Ukraine [3396]
CITES Identification Manual Reference: A-301.011.010.001
Testudo hermanni Gmelin, 1789 II A LR/nt 3240, 3377, 3396, 3436,
3464, 3679
CITES EC Reg. RL References
11
Synonyms: Testudo enriquesi Parenzan, 1932, Testudo hercegovinensis Werner, 1899, Testudo graeca bettai
Lataste, 1881, Testudo graeca hercegovinensis Werner, 1899
E: Hermann's Tortoise, F: Tortue d'Hermann, Tortue d'Hermann, S: Tortuga mediterránea, Tortuga
mediterránea
Albania [3396, 5061, 5161]; Bosnia and Herzegovina [80, 3396]; Bulgaria [3167, 3396]; Croatia [80, 3396]; France
[3279, 3396, 3675, 3676, 6235]: France, Corse; Greece [3396, 3677]; Italy [3212, 3396, 6235]: Sardegna, Sicilia;
Malta (int); Romania [3321, 3396]; Serbia and Montenegro [3396]; Slovenia [80, 3396]; Spain [3396]: Spain
[3458], ?Baleares [6235]; The former Yugoslav Republic of Macedonia [80, 3396]; Turkey [3161, 3396]
CITES Identification Manual Reference: A-301.011.010.002
Testudo horsfieldii Gray, 1844 II B VU 3396, 3436, 3464, 3473
Synonyms: Homopus burnesii Blyth, 1854, Testudo baluchiorum Annandale, 1906, Agrionemys horsfieldii
(Gray, 1844)
E: Afghan Tortoise, Central Asian Tortoise, Four-toed Tortoise, Horsfield's Tortoise, Steppe Tortois